It seems to me that for Darwin the pulsing of evolutionary rates was a strictly vertical phenomenon.
Ernst MayrOn Earth, among millions of lineages or organisms and perhaps 50 billion speciation events, only one led to high intelligence ; this makes me believe its utter improbablity.
Ernst MayrTwo forms or species are sympatric, if they occur together, that is if their areas of distribution overlap or coincide. Two forms (or species) are allapatric, if they do not occur together, that is if they exclude each other geographically. The term allopatric is primarily useful in denoting geographic representatives.
Ernst Mayr