Evolution has no moral direction. An evolutionary understanding of human nature can explain the differing intuitions we have when we are faced with an individual rather than with a mass of people, or with people close to us rather than with those far away, but it does not justify those feelings.

There has been opposition to experimenting on animals for a long time. This opposition has made little headway because experimenters, backed by commercial firms that profit by supplying laboratory animals and equipment, have been able to convince legislators and the public that opposition comes from uninformed fanatics who consider the interests of animals more important than the interests of human beings.

I am not saying that factory farming is the same as the Holocaust or the slave trade, but it's clear that there is an immense amount of suffering in it, and just as we think that the Nazis were wrong to ignore the suffering of their victims, so we are wrong to ignore the sufferings of our victims.

Why...is the hunter who shoots a deer for venison subject to more criticism than the person who buys a ham at the supermarket? Overall, it is probably the intensively reared pig who has suffered more.

Were we incapable of empathy – of putting ourselves in the position of others and seeing that their suffering is like our own – then ethical reasoning would lead nowhere. If emotion without reason is blind, then reason without emotion is impotent.

With the eventual acceptance of Darwin's theory we reach a modern understanding of nature, one which has since then changed in detail rather than in fundamentals. Only those who prefer religious faith to beliefs based on reasoning and evidence can still maintain that the human species is the special darling of the entire universe, or that other animals were created to provide us with food, or that we have divine authority over them, and divine permission to kill them.

Sometimes animals may suffer more because of their more limited understanding. If, for instance, we are taking prisoners in wartime we can explain to them that although they must submit to capture, search, and confinement, they will not otherwise be harmed and will be set free at the conclusion of hostilities. If we capture wild animals, however, we cannot explain that we are not threatening their lives. A wild animal cannot distinguish an attempt to overpower and confine from an attempt to kill; the one causes as much terror as the other.