At a minimum, in explaining evolutionary pathways through time, the constraints imposed by history rise to equal prominence with the immediate advantages of adaptation.
Stephen Jay GouldThe history of life is more adequately represented by a picture of 'punctuated equilibria' than by the notion of phyletic gradualism. The history of evolution is not one of stately unfolding, but a story of homeostatic equilibria, disturbed only 'rarely' (i.e. rather often in the fullness of time) by rapid and episodic events of speciation.
Stephen Jay Gould