The upshot is that most philosophers of biology now hold that biological properties supervene on physical properties (where supervenience is taken to include some kind of "in virtue of" relation), and that fitness and other biological properties are not identical with physical properties.
Elliott SoberDarwin and his successors have railed against the fallacy of confusing the current utility of a trait with the reason the trait evolved. For example, Darwin argued that skull sutures in mammals did not evolve because they facilitate live birth; the sutures were in place well before live birth evolved. Checking the chronological order in which different traits evolved in a lineage is one way to test an adaptive hypothesis; the fact of common ancestry is what makes that checking possible.
Elliott SoberThe big picture, I think, is that common ancestry is evidentially prior to natural selection in Darwin's theory and in contemporary evolutionary biology as well.
Elliott Sober