Paleontologists had long been aware of a seeming contradiction between Darwin's postulate of gradualism, confirmed by the work of population genetics, and the actual findings of paleontology. Following phyletic lines through time seemed to reveal only minimal gradual changes but no clear evidence for any change of a species into a different genus or for the gradual origin of an evolutionary novelty. Anything truly novel always seemed to appear quite abruptly in the fossil record.
Ernst MayrWherever we look at the living biota โฆ discontinuities are overwhelmingly frequentโฆThe discontinuities are even more striking in the fossil record. New species usually appear in the fossil record suddenly, not connected with their ancestors by a series of intermediates.
Ernst MayrEvolution, thus, is merely contingent on certain processes articulated by Darwin: variation and selection. No longer is a fixed object transformed, as in transformational evolution, but an entirely new start is, so to speak, made in every generation.
Ernst MayrLiving in an entirely different physical as well as biotic environment, such a population would have unique opportunities to enter new niches and to select novel adaptive pathways.
Ernst Mayr